I had to move hosts, so the new website for The Theropod Database is
https://theropoddatabase.github.io/ . We're working out some encoding issues for special characters, but in the meantime here are some nomina nuda that have not been reported before.
I'm sure Dalman's new tyrannosaur names have been making the rounds based on his new curriculum vitae at https://www.montana.edu/earthsciences/graduate-program/students/cv/Sebastian_Dalman.html . Here's some informed guesswork as to what they are...
"Bistityrannus" Dalman,
Jasinski, Lucas, Malinzak, Loewen, Fiorillo and Currie, in
progress/review in Dalman, online 2024
?= "Alamotyrannus" Dalman and Lucas, in press in Dalman, 2013 in part
"B. anax" Dalman, Jasinski,
Lucas, Malinzak, Loewen, Fiorillo and Currie, in progress/review in
Dalman, online 2024
?= "Alamotyrannus brinkmani" Dalman and Lucas, in press in Dalman, 2013
in part
Etymologies- Bisti
is Navajo for "large area of shale hills" after the Bisti/De-Na-Zin
Wilderness Area where the specimen was probably found + Latin tyrannus "ruler", common suffix for
tyrannosauroids. Greek anax
"tribal chief or leader".
Alamo after the Ojo Alamo
Formation where the material is from, in turn named for the Ojo Alamo
trading post + Latin tyrannus
"ruler", common suffix for tyrannosauroids. brinkmani probably from vertebrate
paleontologist Donald B. Brinkman.
Late Campanian, Late Cretaceous
De-na-zin Member of Kirtland Formation?, San Juan Basin, New Mexico, US
Material- ?(ACM 7975; intended syntype of "Alamotyrannus
brinkmani") anterior right dentary (~142 mm deep) (Dalman, 2013)
Campanian-Maastrichtian, Late Cretaceous
San Juan Basin, New Mexico, US
?(PMU.R35; = PMU.R85 of Carr and Williamson, 2000 Appendix 1; =
PMU.R1235 of Carr and Williamson, 2000 Fig. 9G-H and Appendix 1;
intended syntype of "Alamotyrannus brinkmani"?)
anterior right dentary (Sullivan and Williamson, 1997)
Comments- ACM 7975 was
discovered in July or August 1924, and figured and briefly described by
Dalman and Lucas (2016) as "Tyrannosauridae indeterminate" despite the
authors listing two characters purportedly differentiating it from
other tyrannosaurids. Namely, "the possession of two foramina
intermandibularis oralis" and "the morphology of the anterior step of
the lingual bar" which "is anterodorsally inclined and has a relatively
short posterior surface (towards the mouth), whereas in other
tyrannosaurids (e.g., Daspletosaurus,
Tarbosaurus, and Tyrannosaurus) the posterior
surface of the step is well-developed and nearly the same length as the
dorsal surface."
Dalman (2013) stated "the many isolated but diagnostic tyrannosaurid
skeletal fossil elements from the Naashoibito Member of the Ojo Alamo
Formation (early Maastrichtian) of northwestern New Mexico (Sullivan et
al. 2005; Jasinski et al. 2011; Dalman and Lucas, in press) provide
evidence for the occurrence of a new taxon of a large tyrannosaurid"
with the in press paper's bibliographic entry naming it "Alamotyrannus
brinkmani." Stuchlik (pers. comm. to Dalman, 7-2018) informs me the
intended holotype was two dentaries, presumedly including ACM 7975 that
was mentioned in Dalman (2013) as "the new Ojo Alamo tyrannosaurid
taxon ACM 7975." Dalman (pers. comm. to Demirjian, 2015) stated the
paper is postponed as more complete remains were discovered, and that
the taxon would receive a different name. A probable explanation
is
that Dalman and Lucas (2016) briefly described and figured ACM 7975 as
a diagnostic tyrannosaurid under study by Dalman but noted that the
previous referral to the Ojo Alamo Formation was due to its discoverer
Loomis using an old, broader definition for the formation.
Geographical and preservational data indicated instead that ACM 7975
was probably from the De-na-zin Member of the Kirtland Formation, so
mixing it with the diagnostic Ojo Alamo elements noted by Dalman (2013)
would make his "Alamotyrannus" concept a chimaera. Thus Dalman
would want to
describe the diagnostic Kirtland dentary and the diagnostic Ojo Alamo
elements as different taxa, which seems to be the plan based on his
2024 online curriculum vitae.
This lists "Dalman, S.G., Jasinski, S.E., Lucas, S.G., Malinzak, D.E.,
Loewen, M.A., Fiorillo, A.R., Currie, P.J. 2024. Bistityrannus anax,
a new tyrannosaurid from the Kirtland Formation (Upper Cretaceous) of
northwestern New Mexico. Cretaceous Research (in review/in progress)"
under Published Research despite being unpublished
as of
9-21-2024. The genus and species are obviously invalid pending
this
publication as his online curriculum vitae is not "issued for the
purpose of providing a public and permanent scientific record" (IZCN
Article 8.1.1), "produced in an edition containing simultaneously
obtainable copies by a method that assures 8.1.3.1. numerous identical
and durable copies (see Article 8.4), or 8.1.3.2. widely accessible
electronic copies with fixed content and layout" (Article 8.1.3), does
not "state the date of publication in the work itself, and" is not
"registered in the Official Register of Zoological Nomenclature
(ZooBank) (see Article 78.2.4) and contain evidence in the work itself
that such registration has occurred" (Articles 8.5.2 and 8.5.3), the
taxa are not "accompanied by a description or definition that states in
words characters that are purported to differentiate the taxon"
(Article 13.1) or "explicitly indicated as intentionally new" (Article
16.1).
Another unpublished entry on that page with identical authorship is "Denazinosaurus sicarius,
a new tyrannosaurid from the Kirtland Formation (De-na-zin Member)
Upper Cretaceous of New Mexico, USA", giving us two proposed new
Kirtland tyrannosaurids with nothing distinguishing them in their
publication titles besides "Denazinosaurus" definitely being from the
De-na-zin Member. While either or neither of these could be
intended
for former "Alamotyrannus" dentary ACM 7975, it's here suggested the
stratigraphic uncertainty behind that specimen's discovery makes
De-Na-Zin unlikely to feature in the article title or genus name.
Thus
ACM 7975 is probably "Bistityrannus", while "Denazinosaurus" would be a
different specimen with a more definite locality. Given most
De-Na-Zin
tyrannosaurid specimens are isolated teeth and postcrania (generally
considered indeterminate in Tyrannosauridae), that ideally Dalman would
want a specimen comparable to ACM 7975 to erect a new contemporaneous
species distinct from it, and that he is describing other
tyrannosaurids based on dentaries (see Fruitland Formation KU
VP-96888), the obvious identity of "Denazinosaurus" would be classic
dentary USNM V 8346 described by Gilmore in 1916 (see entry).
Whether
this logic proves true awaits either publication.
Interestingly, given Dalman originally intended to name "Alamotyrannus"
based on two dentary syntypes including ACM 7975, it implies a second
tyrannosaur dentary plausibly from the Ojo Alamo Formation. While
NMMNH P-7199 is an obvious possibility as the only dentary I know
of confirmed from that formation, it's so poorly preserved that it's
never even been figured. I propose a more likely possibility is
anterior dentary PMU.R35 whose locality can only be specified to San
Juan County, and which seems to have the anterior step morphology
described for ACM 7975- anterodorsally inclined with a short posterior
surface (Sullivan and Williamson, 1997- Fig. 1D). As its
uncertain
stratigraphic placement makes it plausibly from the De-Na-Zin Member,
this may be an additional specimen of "Bistityrannus". Discovered
in
1921 or 1922, this was not published until Sullivan and Williamson's
1997 review of PMU specimens sent by Sternberg, where it is figured in
lateral and medial views as "tyrannosaurid partial right
dentary."
References- Sullivan and
Williamson, 1997. Additions and corrections to Sternberg's San Juan
Basin Collection, Paleontological Museum, University of Uppsala,
Sweden. New Mexico Geological Society Guidebook, 48th Field Conference,
Mesozoic Geology and Paleontology of the Four Corners Region. 255-257.
Carr and Williamson, 2000. A review of Tyrannosauridae (Dinosauria:
Coelurosauria) from New Mexico. In Lucas and Heckert (eds.). Dinosaurs
of New Mexico. New Mexico Museum of Natural History and Science.
Bulletin. 17, 113-146.
Dalman, 2013. New examples of Tyrannosaurus rex from the Lance
Formation of Wyoming, United States. Bulletin of the Peabody Museum of
Natural History. 54(2), 241-254.
Dalman and Lucas, 2016. Frederic Brewster Loomis and the 1924
Amherst
College paleontological expedition to the San Juan Basin, New Mexico.
New Mexico Museum of Natural History and Science Bulletin. 74, 61-66.
Dalman, 2024 online. https://www.montana.edu/earthsciences/graduate-program/students/cv/Sebastian_Dalman.html
Dalman and Lucas, "in press". A new large tyrannosaurid Alamotyrannus
brinkmani, n. gen., n. sp. (Theropoda: Tyrannosauridae), from the
Upper Cretaceous Ojo Alamo Formation (Naashoibito Member), San Juan
Basin, New Mexico. New Mexico Museum of Natural History and Science
Bulletin.
Dalman, Jasinski, Lucas, Malinzak, Loewen, Fiorillo and Currie, in
progress/review a. Denazinosaurus
sicarius,
a new tyrannosaurid from the Kirtland Formation (De-na-zin Member)
Upper Cretaceous of New Mexico, USA. Acta Palaeontologica Polonica.
Dalman, Jasinski, Lucas, Malinzak, Loewen, Fiorillo and Currie, in
progress/review b. Bistityrannus anax,
a new tyrannosaurid from the Kirtland Formation (Upper Cretaceous) of
northwestern New Mexico. Cretaceous Research.
"Denazinosaurus" Dalman,
Jasinski, Lucas, Malinzak, Loewen, Fiorillo and Currie, in
progress/review in Dalman, online 2024
"D. sicarius" Dalman, Jasinski,
Lucas, Malinzak, Loewen, Fiorillo and Currie, in progress/review in
Dalman, online 2024
Etymology- De-Na-Zin is Navajo
for "cranes" and presumedly based on the De-na-zin Member of the
Kirtland Formation which the taxon would be from + Greek sauros "reptile." Latin sicarius "assassin", as it's a
probable hunter.
Late Campanian, Late Cretaceous
head of Hunter Wash / locality 60 of Bauer 1916, De-na-zin Member of
Kirtland Formation, San Juan County, New Mexico, US
Material- ?(USNM V 8346)
incomplete left dentary (tooth row length 354 mm) (Gilmore, 1916)
Comments- Discovered August 20,
1915, this was initially described by Gilmore (1916) as "Deinodon?" and figured in medial
view. He noted "In the number of tooth sockets this jaw agrees
with Dynamosaurus imperosus
Osborn,a [now Tyrannosaurus
rex] but in the general form of the dentary, particularly the
contour of the anterior end, it approaches Albertosaurus4 (Dryptosaurus) most nearly, but as
the dentary of Albertosaurus
has sockets for 15 teeth the presence of 13 in this individual would
appear to show its distinctness." Gilmore further stated "It is
quite
possible that this dentary pertains to the genus Deinodon,
but that can not be determined at this time because the dentary of that
genus is unknown. The identification of this specimen must therefore
await the discovery of additional material." Gilmore (1920)
repeated
most of this verbatim, listing it as "DEINODON, species." this
time.
In a later publication, Gilmore (1935) wrote "Comparison of this bone
directly with a dentary of Gorgosaurus
libratus
Lambe from the Belly River of Canada now shows such close resemblances
in size, shape, and other characteristics down to the smallest details
as to leave little doubt of their being congeneric. Likewise, the
number of alveoli (13) is in agreement with Lambe’s (1917)
determination from a number of specimens that the dentary in this genus
bears 13 or 14 teeth." Lucas et al. (1987) felt the dentary
"almost
certainly pertain[s] to Albertosaurus"
sensu Russell 1970, equating to modern Albertosaurinae. Carr and
Williamson (2000) figured the dentary in lateral, medial and dorsal
views, and stated "Based on personal observation of this and other
tyrannosaurid dentaries, this bone does not appear to be diagnostic at
the specific or generic levels." They noted "There are actually
14
dental alveoli" and concluded "Because the alveolus count of USNM 8346
overlaps with that of A. libratus
and T. rex
and in light of the fact that tooth count is variable in these species,
we consider it best to consider this specimen as indeterminate
Tyrannosauridae." Indeed, 14 dentary alveoli are also present in
some Albertosaurus sarcophagus
(AMNH 5222, TMP 1986.064.0001), Daspletosaurus
torosus (CMN 8506) and Tarbosaurus
(IGM 107/7) specimens.
Dalman's 2024 online curriculum vitae lists "Dalman, S.G. Jasinski,
S.E., Lucas, S.G., Malinzak, D.E., Loewen, M.A., Fiorillo, A.R. and
Currie, P.J. 2024. Denazinosaurus
sicarius,
a new tyrannosaurid from the Kirtland Formation (De-na-zin Member)
Upper Cretaceous of New Mexico, USA. Acta Palaeontologica Polonica (in
review/in progress)" under Published Research despite being unpublished
as of
9-21-2024. The genus and species are obviously invalid pending
this
publication as his online curriculum vitae is not "issued for the
purpose of providing a public and permanent scientific record" (IZCN
Article 8.1.1), "produced in an edition containing simultaneously
obtainable copies by a method that assures 8.1.3.1. numerous identical
and durable copies (see Article 8.4), or 8.1.3.2. widely accessible
electronic copies with fixed content and layout" (Article 8.1.3), does
not "state the date of publication in the work itself, and" is not
"registered in the Official Register of Zoological Nomenclature
(ZooBank) (see Article 78.2.4) and contain evidence in the work itself
that such registration has occurred" (Articles 8.5.2 and 8.5.3), the
taxa are not "accompanied by a description or definition that states in
words characters that are purported to differentiate the taxon"
(Article 13.1) or "explicitly indicated as intentionally new" (Article
16.1).
Another unpublished entry on that page with identical authorship is "Bistityrannus anax, a new
tyrannosaurid from the Kirtland Formation (Upper Cretaceous) of
northwestern New Mexico", giving us two proposed new
Kirtland tyrannosaurids with nothing distinguishing them in their
publication titles besides "Denazinosaurus" definitely being from the
De-na-zin Member. While either or neither of these could be
intended
for former "Alamotyrannus" dentary ACM 7975 (see "Bistityrannus"
entry), it's here suggested the
stratigraphic uncertainty behind that specimen's discovery makes
De-Na-Zin unlikely to feature in the article title or genus name.
Thus
ACM 7975 is probably "Bistityrannus", while "Denazinosaurus" would be a
different specimen with a more definite locality. Given most
De-Na-Zin
tyrannosaurid specimens are isolated teeth and postcrania (generally
considered indeterminate in Tyrannosauridae), that ideally Dalman would
want a specimen comparable to ACM 7975 to erect a new contemporaneous
species distinct from it, and that he is describing other
tyrannosaurids based on dentaries (see Fruitland Formation KU
VP-96888), the obvious identity of "Denazinosaurus" would be classic
dentary USNM V 8346. Whether
this logic proves true awaits either publication.
References- Gilmore, 1916.
Vertebrate faunas of the Ojo Alamo, Kirtland and Fruitland formations.
United States Geological Survey, Professional Paper. 98Q, 279-308.
Gilmore, 1920. Osteology of the carnivorous Dinosauria in the United
States National Museum, with special reference to the genera Antrodemus
(Allosaurus) and Ceratosaurus. Bulletin of the United
States National Museum. 110, 1-154.
Gilmore, 1935. On the Reptilia of the Kirtland Formation of New Mexico,
with descriptions of new species of fossil turtles. Proceedings of the
United States National Museum. 83(2978), 159-188.
Lucas, Mateer, Hunt and O'Neill, 1987. Dinosaurs, the age of the
Fruitland and Kirtland Formations, and the Cretaceous-Tertiary boundary
in the San Juan Basin, New Mexico. In Fassett and Rigby (eds.). The
Cretaceous-Tertiary Boundary in the San Juan and Raton Basins, New
Mexico and Colorado. Geological Society of America Special Paper. 209,
35-50.
Carr and Williamson, 2000. A review of Tyrannosauridae (Dinosauria:
Coelurosauria) from New Mexico. In Lucas and Heckert (eds.). Dinosaurs
of New Mexico. New Mexico Museum of Natural History and Science.
Bulletin. 17, 113-146.
Dalman, 2024 online. https://www.montana.edu/earthsciences/graduate-program/students/cv/Sebastian_Dalman.html
Dalman, Jasinski, Lucas, Malinzak, Loewen, Fiorillo and Currie, in
progress/review a. Denazinosaurus
sicarius,
a new tyrannosaurid from the Kirtland Formation (De-na-zin Member)
Upper Cretaceous of New Mexico, USA. Acta Palaeontologica Polonica.
Dalman, Jasinski, Lucas, Malinzak, Loewen, Fiorillo and Currie, in
progress/review b. Bistityrannus anax,
a new tyrannosaurid from the Kirtland Formation (Upper Cretaceous) of
northwestern New Mexico. Cretaceous Research.
"Erinotonax"
Dalman, Jasinski, Loewen, Malinzak, Currie, Fiorillo and Lucas, in
progress/review in Dalman, online 2024
"E. sabathi" Dalman, Jasinski,
Loewen, Malinzak, Currie, Fiorillo and Lucas, in progress/review in
Dalman, online 2024
Etymology- Perhaps Greek Eri "very" + Greek notos "south" + Greek anax "king"
(suggested by Creisler, pers. comm. 9-2024). sabathi after paleontologist Karol
Sabath.
Late Campanian, Late Cretaceous
Ne-nah-ne-zad Member, Fruitland Formation, San Juan Basin, New Mexico,
US
Material- ?(KU VP-96888) (~10
m) anterior left dentary (140 mm deep) (Dalman and Lucas, 2021)
Comments- Dalman and Lucas
(2021) state "The specimen is being described by us in detail in
another paper as belonging to a new genus and a new species" but
contradictorally identify it as "Tyrannosauridae indet." They
also say "When compared to the dentaries of other tyrannosaurids such
as Albertosaurus, Bistahieversor, Daspletosaurus, Gorgosaurus, Lythronax, Tarbosaurus, Teratophoneus and Tyrannosaurus, the body length of
KUVP-96888 was close to 10 m." This is listed as Tyrannosaurus sp. in the KU online
catalog.
Coincidentally, Dalman's 2024 online curriculum vitae lists "Dalman,
S.G., Jasinski, S.E., Loewen, M.A., Malinzak, D.E., Currie, P.J.,
Fiorillo, A.R., Lucas, S.G. 2024. Erinotonax
sabathi,
a new tyrannosaurid from the Fruitland Formation (Upper Cretaceous) of
New Mexico, USA, insights into the evolution, diversity, and
paleogeography of tyrannosaurids in North America. PeerJ (in review/in
progress)" despite being unpublished
as of
9-21-2024. The genus and species are obviously invalid pending
this
publication as his online curriculum vitae is not "issued for the
purpose of providing a public and permanent scientific record" (IZCN
Article 8.1.1), "produced in an edition containing simultaneously
obtainable copies by a method that assures 8.1.3.1. numerous identical
and durable copies (see Article 8.4), or 8.1.3.2. widely accessible
electronic copies with fixed content and layout" (Article 8.1.3), does
not "state the date of publication in the work itself, and" is not
"registered in the Official Register of Zoological Nomenclature
(ZooBank) (see Article 78.2.4) and contain evidence in the work itself
that such registration has occurred" (Articles 8.5.2 and 8.5.3), the
taxa are not "accompanied by a description or definition that states in
words characters that are purported to differentiate the taxon"
(Article 13.1) or "explicitly indicated as intentionally new" (Article
16.1).
Unless Dalman and Lucas are describing multiple new genera of
tyrannosaurids from the Fruitland Formation, it seems likely
"Erinotonax" is KU VP-96888. Given its fragmentary and
pathological condition, I'm skeptical of any future diagnosis.
References- Dalman and Lucas,
2021. New evidence for cannibalism in tyrannosaurid
dinosaurs from the Upper Cretaceous (Campanian/Maastrichtian) San Juan
Basin of New Mexico. New Mexico Museum of Natural History and Science
Bulletin. 82, 39-56.
Dalman, 2024 online. https://www.montana.edu/earthsciences/graduate-program/students/cv/Sebastian_Dalman.html
Dalman, Jasinski, Loewen, Malinzak, Currie, Fiorillo and Lucas, in
progress/review. Erinotonax sabathi,
a new
tyrannosaurid from the Fruitland Formation (Upper Cretaceous) of New
Mexico, USA, insights into the evolution, diversity, and paleogeography
of tyrannosaurids in North America. PeerJ.
"Atroxicarius" Dalman,
Jasinski, Loewen, Lucas, Malinzak, Fiorillo and Currie, in
progress/review in Dalman, online 2024
"A. eversor" Dalman,
Jasinski, Loewen, Lucas, Malinzak, Fiorillo and Currie, in
progress/review in Dalman, online 2024
Etymology- Latin atrox "fierce" + ? Latin arius "agent of use", except the
'c' is unexplained and atrox
is not a noun. Creisler (pers. comm 9-2024) suggests atrox + Latin cara "head" + Latin suffix -ius, but that should imply cranial
remains which are not at all certain. Greek eversor "destroyer."
Late Maastrichtian, Late Cretaceous
NMMNH L-3961, Naashoibito Member of Ojo Alamo Formation, San Juan
County, New Mexico, US
?(NMMNH P-7199) partial
left dentary, 113 tooth fragments, partial vertebra (Carr and
Williamson, 2000)
Late Maastrichtian, Late Cretaceous
SMP 313b, Naashoibito Member of Ojo Alamo Formation, Hunter Wash,
San Juan County, New Mexico, US
?(SMP VP-1848) incomplete left metatarsal I (~93 mm) (Jasinski,
Sullivan and Lucas, 2011)
Late Maastrichtian, Late Cretaceous
SMP 371, Naashoibito Member of Ojo Alamo Formation, Betonnie Tsosie
Wash, San Juan County, New Mexico, US
?(SMP VP-1113) incomplete right femur (~1 m) (Carr and Williamson, 2000)
Late Maastrichtian, Late Cretaceous
SMP 424b, Naashoibito Member of Ojo Alamo Formation, San Juan Basin,
New Mexico, US
?(SMP VP-2105) incomplete right scapulocoracoid (coracoid ~253 mm
proximodist) (Jasinski, Sullivan and Lucas, 2011)
Late Maastrichtian, Late Cretaceous
Naashoibito Member of Ojo Alamo Formation, San Juan County, New
Mexico, US
?(Ratkevich coll.) right metatarsal IV (~513 mm) (Lehman, 1981)
Late Maastrichtian, Late Cretaceous
Naashoibito Member of Ojo Alamo Formation, San Juan Basin, New Mexico,
US
?(AMNH 5882) right pedal phalanx IV-2 (~143 mm) (Carr and Williamson,
2000)
Comments- Discovered May 7, 1998, NMMNH P-7199 is a partial dentary
associated with tooth fragments and a partial vertebra that all remain
unfigured. Carr and Williamson (2000) refer it "to cf. T. rex
on the basis of its large apical and midheight denticles, 7.5 denticles
per 5 mm and 8.5 denticles per 5 mm, respectively" on the mesial carina
and note the dentary and vertebra "are too weathered and incomplete to
permit identification." Jasinski et al. (2011) suggested however
that
"identification below the family level (Tyrannosauridae) based on
denticle densities is tenuous" so retained it as Tyrannosauridae
indet.. The NMMNH online catalog incorrectly lists this as being
from
the De-na-zin Member of the Kirtland Formation.
Jasinski et al. (2011) notes "The scapulocoracoid (SMP VP-2105, Fig.
7A-B) may be from an adult as it compares readily in size and
morphology to the scapulocoracoids of Tyrannosaurus
rex
(FMNH PR2081; Brochu, 2003, fig. 80). While this indicates the presence
of a large tyrannosaurid in the Naashoibito Member, the specimen cannot
be confidently referred to Tyrannosaurus
rex, but may represent Tyrannosaurus
sp."
Femur SMP VP-1113 is first published as "SMP VP-? femur Naashoibito
Mbr., Kirtland Fm. Tyrannosauridae indet" by Carr and Williamson
(2000), then figured by Lucas and Sullivan (2000) as "Tyrannosauridae,
incomplete femur." Sullivan et al. (2005) identified it as "cf. Daspletosaurus sp., based on
comparison with USNM 10754, a right femur of either Gorgosaurus or Daspletosaurus sp. (labelled as Albertosaurus
sp.) ... from the Dinosaur Park Formation" because "the size and
morphology of SMP VP-1113 closely resembles that of USNM 10754", but
the only stated character is gracility compared to Tyrannosaurus, which is true for
almost all other tyrannosaurids.
Jasinski et al. (2011) figure a metatarsal I (incorrectly called "the
1st phalanx of the 1st metatarsal" on page 226), which along with femur
SMP VP-1113 they state is "from a large tyrannosaurid that seems more
gracile than an adult Tyrannosaurus
rex, although they may represent a juvenile or sub-adult."
Lehman (1981) wrote "single, complete, right fourth metatarsal (on loan
from the collection of Ronald P. Ratkevich of Alamogordo, New Mexico)
is tentatively referred here to Albertosaurus
sp." using the broad concept of the genus popular at the time where it
included Gorgosaurus.
It is figured in anterior, medial and five sectional views.
Carr and Williamson (2000) reported "a large pedal phalanx (AMNH 5882;
Fig. 4A-F) was collected from the Naashoibito Member of the Kirtland
Formation. The collector and date of collection of this specimen are
unknown" and figured it in multiple views as Tyrannosauridae
indet..
Oddly, the AMNH online catalog lists this as being collected by Brown
from the Hell Creek Formation of Snow Creek, Montana, but does
correctly list it as a "2nd phalanx" of Theropoda. Williamson and
Carr
(2005) referred the phalanx to cf. Tyrannosaurus
rex in an abstract.
"Alamotyrannus" into "Atroxicarius"-
Dalman (2013) stated "the many isolated but diagnostic tyrannosaurid
skeletal fossil elements from the Naashoibito Member of the Ojo Alamo
Formation (early Maastrichtian) of northwestern New Mexico (Sullivan et
al. 2005; Jasinski et al. 2011; Dalman and Lucas, in press) provide
evidence for the occurrence of a new taxon of a large tyrannosaurid"
with the in press paper's bibliographic entry naming it "Alamotyrannus
brinkmani." Stuchlik (pers. comm. to Dalman, 7-2018) informs me the
intended holotype was two dentaries, presumedly including ACM 7975 that
was mentioned in Dalman (2013) as "the new Ojo Alamo tyrannosaurid
taxon ACM 7975." Dalman (pers. comm. to Demirjian, 2015) stated the
paper is postponed as more complete remains were discovered, and that
the taxon would receive a different name. A probable explanation
is
that Dalman and Lucas (2016) briefly described and figured ACM 7975 as
a diagnostic tyrannosaurid under study by Dalman but noted that the
previous referral to the Ojo Alamo Formation was due to its discoverer
Loomis using an old, broader definition for the formation.
Geographical and preservational data indicated instead that ACM 7975
was probably from the De-na-zin Member of the Kirtland Formation, so
mixing it with the diagnostic Ojo Alamo elements noted by Dalman (2013)
would make his "Alamotyrannus" concept a chimaera. Thus Dalman
would want to
describe the diagnostic Kirtland dentary and the diagnostic Ojo Alamo
elements as different taxa, which seems to be the plan based on his
2024 online curriculum vitae.
This lists "Dalman, S.G.,
Jasinski, S.E., Loewen, M.A., Lucas, S.G., Malinzak, D.E., Fiorillo,
A.R., and Currie, P.J. 2024. Atroxicarius
eversor,
a new tyrannosaurid from the Ojo Alamo Formation (Upper Cretaceous) of
New Mexico, USA, new insights into the evolution of bistahiversorin
tyrannosaurids in North America. Anatomical Records (in review/in
progress)" under Published Research despite being unpublished as of
9-21-2024 and presumedly referring to The Anatomical Record. This
would then seem to be the new name of the Ojo Alamo tyrannosaurid,
which the title suggests is related to Bistahieversor
in a new tribe "Bistahieversorini", although note Dalman misspelled it
without the first 'e'. The tribe, genus and species are obviously
invalid pending this publication as his online curriculum vitae is not
"issued for the purpose of providing a public and permanent scientific
record" (IZCN Article 8.1.1), "produced in an edition containing
simultaneously obtainable copies by a method that assures 8.1.3.1.
numerous identical and durable copies (see Article 8.4), or 8.1.3.2.
widely accessible electronic copies with fixed content and layout"
(Article 8.1.3), does not "state the date of publication in the work
itself, and" is not "registered in the Official Register of Zoological
Nomenclature (ZooBank) (see Article 78.2.4) and contain evidence in the
work itself that such registration has occurred" (Articles 8.5.2 and
8.5.3), the taxa are not "accompanied by a description or definition
that states in words characters that are purported to differentiate the
taxon" (Article 13.1) or "explicitly indicated as intentionally new"
(Article 16.1) and even a correctly spelled "bistahieversorine" is not
in Latinized form (article 11.7.2) nor is it "accompanied by citation
of the name of the type genus" (Article 16.2). Based on another
Dalman
et al. (in review/progress) citation in Dalman's curriculum vitae, the
De-na-zin tyrannosaurid will be named "Denazinosaurus sicarius" and
preseumedly includes dentary ACM 7975 as the intended holotype.
It's unknown which specimens will be types of "Atroxicarius", but as
Dalman (2013) cited skeletal (not dental, which are generally
undiagnostic) elements described by Sullivan et al. (2005) and Jasinski
et al. (2011), they plausibly include dentary and vertebra NMMNH
P-7199, scapulocoracoid SMP VP-2105, femur SMP VP-1113, metatarsal I
SMP VP-1848, a metatarsal IV from the private Ratkevich collection,
and/or pedal phalanx AMNH 5882.
References- Lehman, 1981. The Alamo Wash local fauna: A new look
at the old Ojo Alamo fauna. In Lucas, Rigby and Kues (eds.). Advances
in San Juan Basin paleontology. University of New Mexico Press. 189-221.
Carr and Williamson, 2000. A review of Tyrannosauridae (Dinosauria:
Coelurosauria) from New Mexico. In Lucas and Heckert (eds.). Dinosaurs
of New Mexico. New Mexico Museum of Natural History and Science.
Bulletin. 17, 113-146.
Lucas and Sullivan, 2000. Stratigraphy and vertebrate biostratigraphy
across the Cretaceous-Tertiary boundary, Betonnie Tsosie Wash, San Juan
Basin, New Mexico. New Mexico Museum of Natural History and Science
Bulletin. 17, 95-103.
Sullivan, Luvas and Braman, 2005. Dinosaurs, pollen, and the
Cretaceous-Tertiary boundary in the San Juan Basin, New Mexico. New
Mexico Geological Society, 56th Field Conference Guidebook, Geology of
the Chama Basin. 56, 395-407.
Williamson and Carr, 2005. Latest Cretaceous tyrannosaurs from the San
Juan Basin, New Mexico. Abstracts of Proceedings from "100 years of Tyrannosaurus
rex, a Symposium." 38.
Jasinski, Sullivan and Lucas, 2011. Taxonomic composition of the Alamo
Wash local fauna from the Upper Cretaceous Ojo Alamo Formation
(Naashoibito Member), San Juan Basin, New Mexico. In Sullivan, Lucas
and Spielmann (eds.). Fossil Record 3. New Mexico Museum of Natural
History and Science Bulletin. 53, 216-271.
Dalman, 2013. New examples of Tyrannosaurus rex from the Lance
Formation of Wyoming, United States. Bulletin of the Peabody Museum of
Natural History. 54(2), 241-254.
Dalman and Lucas, 2016. Frederic Brewster Loomis and the 1924
Amherst College paleontological expedition to the San Juan Basin, New
Mexico. New Mexico Museum of Natural History and Science Bulletin. 74,
61-66.
Dalman, 2024 online. https://www.montana.edu/earthsciences/graduate-program/students/cv/Sebastian_Dalman.html
Dalman and Lucas, "in press". A new large tyrannosaurid Alamotyrannus
brinkmani, n. gen., n. sp. (Theropoda: Tyrannosauridae), from the
Upper Cretaceous Ojo Alamo Formation (Naashoibito Member), San Juan
Basin, New Mexico. New Mexico Museum of Natural History and Science
Bulletin.
Dalman, Jasinski, Loewen, Lucas, Malinzak, Fiorillo and Currie, in
progress/review. Atroxicarius eversor,
a new tyrannosaurid from the Ojo Alamo Formation (Upper Cretaceous) of
New Mexico, USA, new insights into the evolution of bistahiversorin
tyrannosaurids in North America. The Anatomical Record.
Dalman, Jasinski, Lucas, Malinzak, Loewen, Fiorillo and Currie, in
progress/review. Denazinosaurus
sicarius,
a new tyrannosaurid from the Kirtland Formation (De-na-zin Member)
Upper Cretaceous of New Mexico, USA. Acta Palaeontologica Polonica.
But we also have a few other nomina nuda based on early versions of papers.
Fujianvenator Xu, Wang,
Chen, Dong, Lin, Xu, Tang, You, Zhou, Wang, He, Li, Zhang and Zhou, 2023
F. prodigiosus
Xu, Wang, Chen, Dong, Lin, Xu, Tang, You, Zhou, Wang, He, Li, Zhang and
Zhou, 2023
= Fujianvenator "rapidus" Xu,
Wang, Chen, Dong, Lin, Xu, Tang, You, Zhou, Wang, He, Li, Zhang and
Zhou, 2023
Etymology- The apparent early species name "rapidus" is no
doubt
from Latin rapidus "quick",
as the authors state "Fujianvenator
was well adapted for terrestrial locomotion and is likely to have been
capable of running at a high speed."
Comments- Xu et al.'s
(2023)
reporting summary states "The holotype of Fujianvenator rapidus (IVPP
V31985) was discovered in Daxi Basin near Yangyuan Village, Zhenghe
Country, Nanping City, Fujian Province, Southeast China" three times,
making this a likely early name for F. prodigiosus.
Reference- Xu, Wang, Chen,
Dong, Lin, Xu, Tang, You, Zhou, Wang, He, Li, Zhang and Zhou, 2023. A
new avialan theropod from an emerging Jurassic terrestrial fauna.
Nature. 621, 336-343.
Hypnovenator Kubota, Kobayashi and Ikeda, 2024
H. matsubaraetoheorum Kubota, Kobayashi and Ikeda, 2024
= Hypnovenator "sasayamaensis" Kubota, Kobayashi and Ikeda, 2024 online
Etymology- "The specific name, "sasayama",
refers to previous name of a city placed in the eastern region of Hyogo
Prefecture, from where the holotype specimen was collected. It also
refers to an amateur group "Association to Study the Sasayama Group",
to which the two discovers belong."
Comments- The species name "sasayamaensis" was used in the preprint of the article.
References- Kubota, Kobayashi and Ikeda, 2024 online. Early Cretaceous troodontine troodontid (Dinosauria: Theropoda) from
the Ohyamashimo Formation of Japan reveals the early evolution of
Troodontinae. Research Square preprint. DOI: 10.21203/rs.3.rs-4459611/v1
Kubota, Kobayashi and Ikeda, 2024. Early Cretaceous troodontine
troodontid (Dinosauria: Theropoda) from the Ohyamashimo Formation of
Japan reveals the early evolution of Troodontinae. Scientific Reports.
14:16392.
Daspletosaurus wilsoni
Warshaw and Fowler, 2022
= Daspletosaurus "diadematus"
Warshaw, Wilson and Fowler, 2022 online
Etymology- "Diadematus, Latin for crowned, in reference to the novel postorbital horn morphology unique to this species, and its diagnosis as a tyrant dinosaur."
Comments- The initial manuscript called this D. "diadematus", which was replaced by wilsoni after its discoverer John Wilson was removed from the authorship.
References- Warshaw and Fowler, 2022. A transitional species of Daspletosaurus Russell, 1970 from
the Judith River Formation of eastern Montana. PeerJ. 10:e14461.
Warshaw, Wilson and Fowler, 2022 online. A transitional species of Daspletosaurus Russell, 1970 from
the Judith River Formation of eastern Montana. PeerJ reviewing PDF (2022:07:75847:0:1:NEW)
Here's a place where I can post my thoughts on new papers, provide updates on my projects, and post info that will eventually be on my website The Theropod Database - https://theropoddatabase.github.io/ . It will center on theropods, but may delve into other topics as well such as phylogenetics.
Saturday, October 5, 2024
New website for The Theropod Database plus new nomina nuda
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Not sure if it's just me, but there are format issues in the current archive: some special symbols are not properly reported (e.g., in certain Spanish letters: "Ca�ad�n Asfalto").
ReplyDeleteAs I mentioned in my second sentence. Finally figured out how to solve it, but due to a stupid mistake on my part I get to do it letter by letter. Neotheropoda, Dromaeosaurs and Ornithothoraces done, 28 files to go.
ReplyDeleteSorry, I did not understand that sentence meant so.
Delete